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Trilobite*Agnostida *Redlichiida *Corynexochida *Lichida *Phacopida *Proetida *Asaphida *Harpetida *Ptychopariida **Proposed order *Nektaspida Trilobites are extinct arthropods in the class (biology) Trilobita. They appeared in the Cambrian era and flourished throughout the lower Palaeozoic before slowly declining to extinction. The last of the trilobites disappeared in the Permian-Triassic extinction event at the end of the Permian. Trilobites are well-known, possibly the second most famous fossil group after the dinosaurs, and are the most diverse group of animal species preserved in the fossil record, consisting of eight, possibly nine, orders and over 15,000 species. Most were simple, small marine animals that filtered mud to obtain food. ==Physical description== The bodies of trilobites are divided into three parts (''tagmata''): a cephalon (head), composed of the two preoral and first four postoral segments completely fused together; a thorax composed of freely articulating segments; and a pygidium (tail) composed of the last few segments fused together with the telson. In the most primitive trilobites the pygidia are still fairly rudimentary. The thorax is fairly flexible—trilobites are often found curled up like modern sow bugs for protection. [[Image:Trilobite2.jpg|left|thumb|''Asaphus kowalewski'', a trilobite from Russia]] Trilobites had a single pair of preoral antenna (biology)e and otherwise undifferentiated biramous limbs. Each exopodite (walking leg) had six segments, analogous to those of other early arthropods. The first segment also bore a feather-like epipodite, or gill branch, which was used for respiration and swimming. The limbs were covered by lateral projections called ''pleural lobes'', extending outward from a central ''axial lobe''. Contrary to popular belief, it is this longitudinal tripartite division into left and right pleural lobes and a central axial lobe that gives trilobites their name, ''not'' the division into cephalon, thorax and pygidium. Trilobites were only armored on top, but still had a fairly heavy exoskeleton. In molting it generally split between the head and thorax, which is why so many trilobite fossils are missing one or the other: many trilobite fossils are actually molts, rather than dead individuals. In most groups there were two facial sutures on the cheeks to make shedding easier. The cheeks usually also supported a pair of crescent-shaped compound eyes, which were surprisingly advanced in some species. In fact, trilobites were the first animals to evolve true eyes, about 543 million years ago; the evolutionary appearance of eyes has been postulated as a trigger for the Cambrian Explosion. Some trilobites like those of the order Lichida evolved elaborate spiny forms, particularly during the Devonian period. (For example, such specimens are found in the Devonian Hamar Laghdad formation of Alnif in Morocco). This could possibly have been a defensive response to the evolutionary appearance of fishes. According to "New Scientist" magazine (May 2005), "some... trilobites... had horns on their heads similar to those of modern beetles." Based on the size, location, and shape of the horns, Rob Knell, a biologist at Queen Mary University of London and Richard Fortey of London's Natural History Museum, concluded that the most likely use of the horns was combat for mates, making trilobites the earliest example of this behavior. While this study only covered the raphiophorid family, the conclusions can be applied to other groups as well, such as the ''Walliserops trifurcates'' Trilobites range in length from one millimetre to 70 cm (1/25 inch to 20 inches), with a typical size range of two to seven centimetres (1 to 3 1/2 inches). ==Sensory organs== Many trilobites had eyes and they had antenna (biology)e that perhaps were used for taste and smell. Some trilobites were blind, probably living too deep in the sea for light to reach them. Others, such as ''Phacops rana'', had eyes that were quite large. The eyes of trilobites were made of calcite (calcium carbonate, CaCO3). Pure forms of calcite are transparent, and the trilobites used clear calcite crystals to form the lenses of their eyes. In this, they differ from most other arthropods, which have soft eyes. The trilobite eyes were typically compound, with each lens being an elongated prism. The number of lenses in such an eye varies, however: some trilobites had only one so they could only distinguish between light and dark, and some had thousands of lenses in one eye. In these compound eyes, the lenses are arranged hexagonally. A living species with similar eyes is the brittle star ''Ophiocoma wendtii''. === Holochroal eyes === Holochroal eyes had a great number of (tiny) lenses (up to over 15000), and are found in all orders of trilobite. These lenses were packed closely together (hexagonally) and touched eachother. A single cornea covered all lenses. These eyes had no sclera, the white layer covering most current eyes. === Schizochroal eyes === Schizochroal eyes typically had less (and larger) lenses (to around 700), and are found only in Phacopida. The lenses were separate, with each lens having an individual cornea which extended into a rather large sclera. === Abathochroal eyes === Abathochroal eyes had few (and small) lenses (to around 70), and are found only in agnostida. Each lens was separate and had an individual cornea. The sclera was separate from the cornea, and did not run as deep as the sclera in schizochroal eyes. ==Development== An egg hatched to give a tiny larva called a ''protaspid'', in which only the fused segments of the cephalon are present. Subsequent thoracic segments were added behind the cephalon in successive molts during an intermediate stage called ''meraspid'', until finally the adult number of segments was reached, at which point the animal is called a ''holaspid''. Trilobite larvae are reasonably well known and provide an important aid in their classification. ==Extinction== The exact reason for the extinction of the trilobites is not clear, although it would seem to be no coincidence that their numbers began to decrease with the arrival of the first sharks and other early fishes in the Silurian and Devonian periods. Trilobites may have provided a rich source of food for these new arrivals. Their relatively low numbers and diversity at the end of the Permian no doubt contributed to their extinction during that great mass extinction event. ==Fossil distribution== Trilobites appear to have been exclusively marine organisms since the fossilized remains of trilobites are always found in rock containing fossils of other salt-water animals such as brachiopods, crinoids, and coral, and they are found in a range of environments from extremely shallow water to very deep water. The tracks left behind by trilobites crawling on the sea floor are occasionally preserved as trace fossils. Trilobite fossils are found worldwide, with many thousands of known species. Because they evolved rapidly, trilobites serve as excellent index fossils, enabling geologists to date the age of the rocks in which they are found. They were among the first fossils to attract widespread attention, and new species are being discovered every year. Native Americans, recognizing that trilobites were water creatures, had a name for them which means "little water bug in the rocks". A famous location for trilobite fossils in the United Kingdom is Wren's Nest, Dudley in the West Midlands (county), where ''Calymene blumenbachi'' is found in the Silurian Wenlock Limestone formation. This trilobite is featured on the town's coat of arms and was named the "Dudley locust" or "Dudley bug" by quarrymen who once worked many of the now abandoned limestone quarries. Other trilobites found there include ''Dalmanites'', ''Trimerus'' and ''Bumastus''. == References == * ''Trilobite!'' - Richard Fortey (ISBN 0002570122) * Riccardo Levi-Setti. ''Trilobites''. University of Chicago Press, 1993. * [http://www.trilobites.info/ A Guide to the Orders of Trilobite] by Sam Gon III - an excellent, well-researched site with information covering trilobites from all angles. Includes many line drawings and photographs. *[http://www.newscientist.com/channel/life/mg18625015.100/ Earliest combatants in sexual contests revealed] from "New Scientist" magazine. == See also == * Prehistoric life Fossils Prehistoric arthropods TrilobiteWith respect to the limbs, is biramous the right word to use? I know it has two branches, but they are not homologous at all to those of crustacean limbs. AFAIK. The (1959)Treatise devotes a couple of pages to 'Biramous Appendages' and I've seen the term in a fair number of other places. ---- Are we allowed to reference other sites? Probably we should have a link to Sam Gon's trilobite page? http://www.aloha.net/~smgon/ ---- Yes we are, just as we can reference books. I'll add that. --- N.B. I followed the link to "Trilobite Orders" and noticed a note saying that the site will be moving soon to a new domain. So I thought I'd record some notes so it can be found again, next time someone looks and finds the link broken: The top of the page reads A Guide to the Orders of Trilobites A site devoted to understanding trilobites created and maintained by Sam Gon III so try searching for that. Also contains "Trilobite of the Month". Added text from an article I originally wrote in 1998 and published on the Web. User:Dlloyd 21:33, 27 Jul 2004 (UTC) Portions of this text are : "Copyright 1995-1997 The Fossil Company Ltd. 1997-1999 The British Fossil Company Inc. and licensed by the owner under the terms of the Wikipedia copyright." Please contact me if you need further clarification on this. User:Dlloyd 00:50, 30 Jul 2004 (UTC) == Trilobite Eyes == ''Trilobites had unique eyes, which were made of calcite (calcium carbonate, CaCO3).'' How would anyone ever know what trilobite eyes were made of? I thought all we know about trilobites comes from fossils that have completely replaced the original substances of the eyes (whatever it might be) with rock. --User:DavidCary 01:15, 6 Nov 2004 (UTC) I'll try to look into this and see if I can answer your question. --User:DanielCD 21:38, 8 Nov 2004 (UTC) The answer is, I believe, that calcite is a fairly stable material, and so under the conditions that many trilobites were preserved, there would have been no replacement of the calcite by other minerals. The optical properties of calcite in the eye is so well preserved that you can "see" through the lenses and test the visual acuity of the original creatures. Hope this helps. --User:DanielCD 21:48, 9 Nov 2004 (UTC) =="Native Americans"== ''"The Native Americans had a name for trilobite which means "little water bug in the rocks"a name which demonstrates extraordinary zoological acuity."'' This is like saying "The Asians had a name for..." --it's meaningless. Doubtless a Western North American tribe is meant. The "zoological acuity" would depend on the phrase being ''independent'' of a paleontologist saying, "and what would you call these?" Not to denigrate the zoological and botanical acuity of any of the "First Peoples." --User:Wetman 17:18, 11 Nov 2004 (UTC) Well, fix it then. --User:DanielCD 21:43, 11 Nov 2004 (UTC) == Trilobite Pictures == I was reading the article for the word "Fossil" and noticed that there is a very nice picture of a fossilized trilobite, complete with its eyestalks still attached. I think that this image should replace one of the ones on the page for trilobites. What do you all think? Here is the picture I am talking about: http://en.wikipedia.org/wiki/Image:Trilobite2.jpg == Trilobite Pictures II == I added the pic mentioned above, but also changed the pic in the taxobox. The reason being that this one is really clear and shows the tripartate structure really well. Also, the pic of the trilobite in Death Valley got bumped due to space. I really didn't think it was that great anyway. If anyone wants it back, we can pull it from the history and restore it. --User:DanielCD 21:11, 19 Apr 2005 (UTC) Trilobite
== turnout in EU const. vote overview == Hi Trilobite, I couldn't find how to modify that upper-right table with the referendum overview for the proposed EU constitution... could you point me how to do it? (or add the turnouts yourself if you wish?) Thanks again! --User:Jordi Burguet Castell 23:20, 1 Jun 2005 (UTC) :Replied on talk page. — User:Trilobite 23:41, 1 Jun 2005 (UTC) ::Thanks much, I didn't know about templates indeed! --User:Jordi Burguet Castell 00:06, 2 Jun 2005 (UTC) ==Robert McCartney== Hi. I was just wondering if you are going to update the McCartney article, as it has not been updated since the family were in the USA? User:Fergananim :Hi, it's probably not a good idea to leave messages in my talk page archives; I only just found that one! I may update it some time, but there is a fair bit of work to be done and I am on a partial wikibreak at the moment. You could have a go yourself, of course. I should think just about all the news reports you'd need are available on the web. I'll have a look at it myself eventually and see what needs doing. Sorry not to be of more assistance. I've been meaning to get round to sorting it out for a while now (always the way on Wikipedia). Cheers. — User:Trilobite 03:02, 13 Jun 2005 (UTC) == Thank you for your support == Thank you for supporting my candidacy for administrator. User:Kelly Martin 14:43, Jun 10, 2005 (UTC) ==W. Mark Felt== Salve! I went and nominated W. Mark Felt as a WP:FAC. As you commented on it on the talk page, I'd appreciate your comments at Wikipedia:Featured article candidates/W. Mark Felt. User:PedanticallySpeaking 14:54, Jun 17, 2005 (UTC) :Thanks for the note. I remember trying to keep the article in something like a reasonable condition on the day the news broke and edits were coming in thick and fast. I am no Watergate expert and so probably not qualified to make a call on the nomination, but I'll keep an eye on it and comment if I feel something needs mentioning. — User:Trilobite 23:08, 17 Jun 2005 (UTC) == Ubuntu == Thanks for running to the rescue at Ubuntu. I've always been removing those extlinks on sight, but I never came up with the idea of adding a hidden comment for those illiterate Ubuntu Linux fans. In fact, I was having fun inventing new 'baffled and thunderstruck' edit summaries from time to time :P . Kind regards, — User:Mark Dingemanse User Talk:Mark Dingemanse 19:27, 20 Jun 2005 (UTC) :Thanks and sorry to spoil your fun with the edit summaries! I wonder if someone will now fail to notice the comments and add Linux in again. Judging by some of the comments that regularly appear on Talk:Main Page despite the huge notice at the top telling everyone it's not a general contact point it would not surprise me greatly. — User:Trilobite 20:12, 20 Jun 2005 (UTC) == Thanks == Trilobite: thanks for the advice, when I have anything worth archiving (like you) I will do that, isn't all in history anyway? User:Wiki alf 14:22, 22 Jun 2005 (UTC) My replies are on my talk. User:Wiki alf 15:31, 22 Jun 2005 (UTC) :Thanks, I saw them. — User:Trilobite 15:34, 22 Jun 2005 (UTC) Trilobite: I would like to prove I have ox.chem.ac.uk's permission for an article, who would I forward their email to for this purpose? User:Wiki alf 20:40, 22 Jun 2005 (UTC) :You would probably need to paste the email in question on the talk page of the article. By the way, it's important to note that permission to reproduce copyrighted material ''on this website'' is not good enough, as Wikipedia is free content that anyone can reproduce and modify as they wish. I assume you got them to release it under the GNU Free Documentation License, in which case everything is fine from a legal angle. Often it's not a good idea to add content to Wikipedia that was not written specifically for us, as it may need extensive work before it reads like a proper Wikipedia article. I don't know about your individual circumstances, however. If you're trying to clear up a dispute surrounding an article that was flagged as a copyvio then putting the relevant permission on the talk page should clear things up. Hope this helps. — User:Trilobite 20:54, 22 Jun 2005 (UTC) :OK already pasted the email into relevant page (glad I got something right) didn't specifically ask for GFDL but as it is virtually only the books references and research topic bullets would this be a big issue, as I'm pretty sure that was already in public domain? Maybe you'd take a look at their site and then the article and feedback any concerns. I would be surprised if his entry survives however so might not matter anyway. ox.chem website is [http://www.chem.ox.ac.uk/researchguide/dmhodgson.html] and the article is David Hodgson (chemist)/Temp.User:Wiki alf 21:04, 22 Jun 2005 (UTC) ::With such a short entry consisting mainly of things like book references, copying it without permission would probably not cause a huge problem, but we need permission nonetheless. Unless it's several decades old, everything published that doesn't consist of simple facts (like a list of cities the Olympics has been held in, or something), and that hasn't been explicitly released into the public domain by its copyright holder should be considered to be protected by copyright law and not inserted into Wikipedia. Some people wrongly assume that a website without a copyright notice is public domain, but anyway, since the one you linked to explicitly claims copyright for the University of Oxford at the bottom we definitely couldn't have used it without permission. It's not as if these things are ever likely to end in a court case or something, it's just that to avoid any question of copyright violation Wikipedia has to take a tough line on such matters. I try to avoid getting into these copyright disputes so I'll leave it for people who know more than me about these things to sort it out. I should think you've cleared it up by putting the note on the talk page. Cheers. — User:Trilobite 21:22, 22 Jun 2005 (UTC)d :::I am indebted, many thanks. User:Wiki alf 23:29, 22 Jun 2005 (UTC) TrilobiteI have been a reader of Wikipedia since early 2003, an editor and contributor since May 2004, and an administrator since April 2005. I don't expect to have time to edit much over the next few weeks and months, but I'll be back eventually. I am in the process of documenting and developing some User:Trilobite/Tools, which are mainly concerned with speeding up common tasks or modifying the interface, and which others may find helpful. This is very much a work in progress. In case you were wondering, my username has no special significance, and I can't even remember why I chose it. As I recall I just plucked it out of thin air when I registered an account. At the time I didn't plan on staying around and making thousands of edits, but Wikipedia is extremely addictive. ---- Things to keep an eye on: *Wikipedia:Wikipedia Signpost *Wikipedia:Village pump (all) *Wikipedia:Administrators' noticeboard *Wikipedia:Requests for adminship *[http://mail.wikipedia.org/ Wikimedia mailing lists] **[http://mail.wikipedia.org/pipermail/foundation-l/-/thread.html Wikimedia Foundation] **[http://mail.wikipedia.org/pipermail/wikipedia-l/-/thread.html Global Wikipedia] **[http://mail.wikimedia.org/pipermail/wikien-l/-/thread.html English Wikipedia] ---- ---- {| cellspacing="1" cellpadding="1" style="background: #F1F1DE; border: 1px solid #CC9; padding: 5px;" |- style="text-align: center;" | style="background: #CCC;" |Multi-licensed into the public domain |- | colspan="3" |''I agree to multi-license my eligible text contributions, unless otherwise stated, under the [http://www.gnu.org/copyleft/fdl.html GFDL] and into the public domain. Please be aware that other contributors might not do the same, so if you want to use my contributions in the public domain, please check the Wikipedia:Multi-licensing. This does not include my user pages. The copyright status of images I have uploaded is indicated on their individual image description pages.'' |} {| width="80%" align="center" style="text-align:center; border:1px solid #ffc9c9; background-color:#FFFFF3;" |- padding:1em;padding-top:0.5em;" | |style="font-size: 85%"|This is a Wikipedia user page. If you find this page on any site other than [http://www.wikipedia.org/ Wikipedia], you are viewing a mirror site. Be aware that the page may be outdated, and that the user this page belongs to has no personal affiliation with any site containing Wikipedia content other than Wikipedia itself. Wikipedia mirrors are generally poor imitations run by people who are only interested in showing you advertising. Often they fail to comply with the terms of Wikipedia's copyleft licence or attempt to hide the origin of their content. The original version of this page is located at http://en.wikipedia.org/wiki/User:Trilobite. |} __NOTOC__ __NOEDITSECTION__ Public domain license See other meanings of words starting from letter: TTA | TB | TC | TD | TE | TF | TG | TH | TI | TJ | TK | TL | TŁ | TM | TN | TO | TP | TR | TS | TU | TW | TX | TY | TZ |Words begining with Trilobite: Trilobite Trilobite Trilobite Trilobite Trilobite/Archive1 Trilobite/Archive2 Trilobite/Archive3 Trilobite/County_Durham Trilobite/draft1 Trilobite/draft2 Trilobite/draft3 Trilobite/draft4 Trilobite/draft5 Trilobite/MEP_script Trilobite/monobook.css Trilobite/monobook.js Trilobite/sandbox Trilobite/Tools Trilobites |
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